TYPE: Costa Rica. Puntarenas: Aguirre, road between Quepos and San Marcos de Terrazú, 9º35’N 84º06’W, 830 m, premontane wet forest, epiphytic in secondary vegetation along the roadside, 16 February 2002, F. Pupulin, D. Castelfranco, H. Montealegre & A.C. Rodríguez 3517 (holotype, USJ; isotypes, USJ-79001, USJ, USJ-Spirit, CR; illustration of type).